Six months have elapsed since the successful Battle of Britannia and the defeat of the Neuroi invaders in Gallia. Yoshika Miyafuji, a distinguished member of the 501st Joint Fighter Wing, has returned to Fuso and finished middle school. Nonetheless, her mission isn't done yet. She receives a letter sent by her deceased father with blueprints for a cutting-edge Striker Unit he had been creating before his death. The unit, made specifically for Yoshika, may have the power to harness her extraordinary magical abilities. In the meantime, a new danger is emerging in Europe. A Neuroi nest with immense size and force has appeared over Venezia, destroying local Witch forces and devouring the northern part of the nation. To add to the chaos, a new kind of Neuroi with a human-like form is attempting to make contact with people. Yoshika must not abandon her comrades on the front lines and return to the ravaged continent to face this new challenge.