A dozen millennia have passed since the events described in Genesis Aquarion. Today, humans reside on Vega, a star that is constantly under attack from trans-dimensional creatures called Abductors, who hail from the neighboring star, Altair. As a countermeasure, the organization called Neo-DEAVA was formed, whose members pilot highly advanced mecha suits called Aquaria. The genders are strictly separated and boys and girls are prohibited from interacting, let alone fighting together on the battlefield. But everything changes when an Abductor mecha suit of unprecedented sophistication enters the fray. Two teenagers, Mikono and Amata, are unwittingly caught up in the conflict. Amata commits an unspeakable act when he summons an Aquaria and forms a Forbidden Union between a male and female mecha suit. Neo-DEAVA is left in shock, and the implications of Amata’s violations reach further than he could have ever imagined. How was he able to summon and control an Aquaria? Where did he learn about the Forbidden Union? And why is Mikono also capable of piloting a mecha suit of this kind?